aniPronunciation: /ˈɑːni/Definition of ani noun (plural anis) a glossy black long-tailed bird of the cuckoo family, with a large deep bill, found in Central and South America. Genus Crotophaga, family Cuculidae: three species Origin: early 19th century: from Spanish ani, Portuguese anum, from Tupi anũani in other Oxford dictionaries Definition of ani in the US English dictionary