epicuticle Syllabification: (ep·i·cu·ti·cle)Pronunciation: /ˌepiˈkyo͞otikəl/Definition of epicuticle noun Botany & Zoology the thin, waxy, protective outer layer covering the surfaces of some plants, fungi, insects, and other arthropods. Derivatives epicuticularPronunciation: /-kyo͞oˈtikyələr/ adjective epicuticle in other Oxford dictionaries Definition of epicuticle in the British & World English dictionary