crepuscular Syllabification: (cre·pus·cu·lar)Pronunciation: /krəˈpəskyələr/Definition of crepuscular adjective of, resembling, or relating to twilight. Zoology (of an animal) appearing or active in twilight. Origin: mid 17th century: from Latin crepusculum 'twilight' + -ar1crepuscular in other Oxford dictionaries Definition of crepuscular in the British & World English dictionary