epsilonPronunciation: /ˈɛpsɪlɒn, ɛpˈsʌɪlɒn/Definition of epsilon noun the fifth letter of the Greek alphabet (Ε, ε), transliterated as ‘e’. [as modifier] denoting the fifth in a series of items, categories, etc.: epsilon and gamma globins diverged about 100 million years ago (Epsilon) [followed by Latin genitive] Astronomy the fifth star in a constellation:Epsilon Carinae symbol (ε) permittivity. Origin:Greek, 'bare or simple E', from psilos 'bare'epsilon in other Oxford dictionaries Definition of epsilon in the US English dictionary