crepuscularPronunciation: /krɪˈpʌskjʊlə, krɛ-/Definition of crepuscular adjective resembling or relating to twilight. Zoology (of an animal) appearing or active in twilight. Origin: mid 17th century: from Latin crepusculum 'twilight' + -ar1crepuscular in other Oxford dictionaries Definition of crepuscular in the US English dictionary