MalacostracaPronunciation: /ˌmaləˈkɒstrəkə/Definition of Malacostraca noun Zoology a large class of crustaceans which includes crabs, shrimps, lobsters, isopods, and amphipods. They have compound eyes, which are typically on stalks. Origin:modern Latin (plural), from malaco- 'soft' + Greek ostrakon 'shell'Malacostraca in other Oxford dictionaries Definition of Malacostraca in the US English dictionary