speciation Syllabification: (spe·ci·a·tion)Pronunciation: /ˌspēSHēˈāSHən, ˌspēsē-/Definition of speciation noun Biology the formation of new and distinct species in the course of evolution. Derivatives speciatePronunciation: /ˈspēSHēˌāt, spēsē-/ verb speciation in other Oxford dictionaries Definition of speciation in the British & World English dictionary