phloem Syllabification: (phlo·em)Pronunciation: /ˈflōˌem/Definition of phloem noun Botany the vascular tissue in plants that conducts sugars and other metabolic products downward from the leaves. Origin: late 19th century: from Greek phloos 'bark' + the passive suffix -ēmaphloem in other Oxford dictionaries Definition of phloem in the British & World English dictionary